This title exposes and evaluates a set of conceptual disputes concerning what we might mean by culture, and how we should go about accounting for it. Its particular focus is a set of evolutionary approaches to the genesis of the human capacity for culture, to subsequent cultural change, and to the ways in which genetic and cultural change interact, or 'co-evolve'. The book as a whole argues that there is little realistic hope that the social sciences might become unified around an evolutionary synthesis. Instead the defence of evolutionary approaches to culture must be more modest in scope.